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Basic Characteristics of Mutations
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Mutation Site
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P323L |
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Mutation Site Sentence
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SARS-CoV-2 variants with NSP12 P323L/G671S mutations display enhanced virus replication in ferret upper airways and higher transmissibility. |
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Mutation Level
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Amino acid level |
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Mutation Type
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Nonsynonymous substitution |
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Gene/Protein/Region
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NSP12 |
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Standardized Encoding Gene
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ORF1b
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Genotype/Subtype
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Alpha;Beta;Delta;Omicron(BA.1;BA.5;BA.2.75;XBB) |
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Viral Reference
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EPI_ISL_406801
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Functional Impact and Mechanisms
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Disease
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Cell line
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Immune
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- |
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Target Gene
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-
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Clinical and Epidemiological Correlations
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Clinical Information
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- |
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Treatment
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- |
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Location
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- |
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Literature Information
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PMID
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37676771
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Title
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SARS-CoV-2 variants with NSP12 P323L/G671S mutations display enhanced virus replication in ferret upper airways and higher transmissibility
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Author
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Kim SM,Kim EH,Casel MAB,Kim YI,Sun R,Kwak MJ,Yoo JS,Yu M,Yu KM,Jang SG,Rollon R,Choi JH,Gil J,Eun K,Kim H,Ensser A,Hwang J,Song MS,Kim MH,Jung JU,Choi YK
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Journal
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Cell reports
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Journal Info
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2023 Sep 26;42(9):113077
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Abstract
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With the emergence of multiple predominant SARS-CoV-2 variants, it becomes important to have a comprehensive assessment of their viral fitness and transmissibility. Here, we demonstrate that natural temperature differences between the upper (33 degrees C) and lower (37 degrees C) respiratory tract have profound effects on SARS-CoV-2 replication and transmissibility. Specifically, SARS-CoV-2 variants containing the NSP12 mutations P323L or P323L/G671S exhibit enhanced RNA-dependent RNA polymerase (RdRp) activity at 33 degrees C compared with 37 degrees C and high transmissibility. Molecular dynamics simulations and microscale thermophoresis demonstrate that the NSP12 P323L and P323L/G671S mutations stabilize the NSP12-NSP7-NSP8 complex through hydrophobic effects, leading to increased viral RdRp activity. Furthermore, competitive transmissibility assay reveals that reverse genetic (RG)-P323L or RG-P323L/G671S NSP12 outcompetes RG-WT (wild-type) NSP12 for replication in the upper respiratory tract, allowing markedly rapid transmissibility. This suggests that NSP12 P323L or P323L/G671S mutation of SARS-CoV-2 is associated with increased RdRp complex stability and enzymatic activity, promoting efficient transmissibility.
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Sequence Data
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GISAID
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